Pyrenodesmia alociza (A. Massal.) Arnold

Flora, 67: 310, 1884. Basionym: Biatorina alociza A. Massal. - Symmicta Lich.: 42, 1855.
Synonyms: Caloplaca alociza (A. Massal.) Mig.; Lecaniella alociza (A. Massal.) Jatta; Lecanora variabilis f. ecrustacea Nyl.; Placodium variabile var. ecrustaceum (Nyl.) Nyl.; Sporoblastia alocyza (A. Massal.) Trevis.
Distribution: N - VG (Crisafulli & al. 2006, Piervittori & al. 2006, Tretiach & Muggia 2006), Frl, Ven (Lazzarin 2000b, Tretiach & al. 2003, Nascimbene & Caniglia 2003c, Tretiach & Muggia 2006, Muggia & al. 2007, Thor & Nascimbene 2007), TAA (Spitale & Nascimbene 2012), Emil. C - Tosc (TSB 37172), Marc (Nimis & Tretiach 1999), Umb (Nimis & Tretiach 1999, Ravera & al. 2006, Panfili 2007), Laz (Nimis & Tretiach 2004), Abr (Nimis & Tretiach 1999), Mol (Nimis & Tretiach 1999, Caporale & al. 2008, Genovesi & Ravera 2014), Sar. S - Camp (Aprile & al. 2003b, Nimis & Tretiach 2004, Garofalo & al. 2010), Pugl (Nimis & Tretiach 1999, Tretiach & Muggia 2006), Bas (Nimis & Tretiach 1999), Cal (Puntillo 1996), Si (Nimis & al. 1994, Grillo 1998, Grillo & Caniglia 2004, Caniglia & Grillo 2005, 2006).
Description: Thallus crustose, endolithic or (rarely) very thinly epilithic, white to grey, sometimes with a violet hue, often delimited by a dark prothallus. Apothecia 0.2-0.8(-1) mm across, often slightly immersed in pits in the rock, with a flat to slightly convex, not pruinose, black disc and a thin, finally excluded proper margin. Proper exciple brown in outer part, colourless to brownish within, the pigmented parts K+ violet; epithecium grey to brownish (in thick sections), K+ violet; hymenium colourless, with small crystals; apical cells of paraphyses 2-5 μm wide; hypothecium colourless. Asci 8-spored, clavate to broadly clavate, Teloschistes type. Ascospores 1-septate, polarilocular, hyaline, ellipsoid, (11-)15-18(-21) x (6-)7-8(-11) μm, the septum 2.5-3 μm thick. Photobiont chlorococcoid. Spot tests: thallus K-, C-, KC-, P-; pigmented parts of apothecia K+ violet and sometimes N+ pale violet. Chemistry: no lichen substances in thallus, Sedifolia-grey pigment in apothecia.
Note: on hard limestones and dolomite on sunny, exposed cliffs, with a wide altitudinal range; in Italy this is one of the most frequent species of Pyrenodesmia. For further details see Muggia & al. (2007).
Growth form: Crustose endolithic
Substrata: rocks
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly sexual

Commonnes-rarity: (info)

Alpine belt: common
Subalpine belt: very common
Oromediterranean belt: extremely common
Montane belt: extremely common
Submediterranean belt: common
Padanian area: rare
Humid submediterranean belt: common
Humid mediterranean belt: rather rare
Dry mediterranean belt: rare


pH of the substrata:

(info)

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5

Solar irradiation:

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Aridity:

(info)

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Eutrophication:

(info)

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5

Poleotolerance:

(info)

0

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Altitudinal distribution:

(info)

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Felix Schumm – CC BY-SA 4.0
[16492], Germany, Baden-Württemberg, Rems-Murr-Kreis, Stetten/Rems, Haldenbachtal, auf Mörtel an schattiger Mauer im Wald, 311 m, 48,7764° N, 9,998° E, leg. et det. Schumm 12.05.2010.


P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (15613)
2001/12/18


Domenico Puntillo; Owner: Domenico Puntillo
Italy


Andrea Moro; Owner: Department of Life Sciences, University of Triedte
Italy, Friuli Venezia Giulia, Trieste, Trieste Karst, Rocca di Monrupino
20/02/2017


Andrea Moro; Owner: Department of Life Sciences, University of Triedte
Italy, Friuli Venezia Giulia, Trieste, Trieste Karst, Rocca di Monrupino
20/02/2017


Andrea Moro; Owner: Department of Life Sciences, University of Triedte
Italy, Friuli Venezia Giulia, Trieste, Trieste Karst, Rocca di Monrupino
20/02/2017


Andrea Moro; Owner: Department of Life Sciences, University of Triedte
Italy, Friuli Venezia Giulia, Trieste, Trieste Karst, Rocca di Monrupino
20/02/2017


Felix Schumm – CC BY-SA 4.0
[16492], Germany, Baden-Württemberg, Rems-Murr-Kreis, Stetten/Rems, Haldenbachtal, auf Mörtel an schattiger Mauer im Wald, 311 m, 48,7764° N, 9,998° E, leg. et det. Schumm 12.05.2010.


Felix Schumm – CC BY-SA 4.0
[16492], Germany, Baden-Württemberg, Rems-Murr-Kreis, Stetten/Rems, Haldenbachtal, auf Mörtel an schattiger Mauer im Wald, 311 m, 48,7764° N, 9,998° E, leg. et det. Schumm 12.05.2010.


Felix Schumm – CC BY-SA 4.0
[16492], Germany, Baden-Württemberg, Rems-Murr-Kreis, Stetten/Rems, Haldenbachtal, auf Mörtel an schattiger Mauer im Wald, 311 m, 48,7764° N, 9,998° E, leg. et det. Schumm 12.05.2010.


Felix Schumm CC BY-SA 4.0
[16492], Germany, Baden-Württemberg, Rems-Murr-Kreis, Stetten/Rems, Haldenbachtal, auf Mörtel an schattiger Mauer im Wald, 311 m, 48,7764° N, 9,998° E, leg. et det. Schumm 12.05.2010.


Felix Schumm CC BY-SA 4.0
[16492], Germany, Baden-Württemberg, Rems-Murr-Kreis, Stetten/Rems, Haldenbachtal, auf Mörtel an schattiger Mauer im Wald, 311 m, 48,7764° N, 9,998° E, leg. et det. Schumm 12.05.2010.


Felix Schumm CC BY-SA 4.0
[16492], Germany, Baden-Württemberg, Rems-Murr-Kreis, Stetten/Rems, Haldenbachtal, auf Mörtel an schattiger Mauer im Wald, 311 m, 48,7764° N, 9,998° E, leg. et det. Schumm 12.05.2010.


Felix Schumm CC BY-SA 4.0
[16492], Germany, Baden-Württemberg, Rems-Murr-Kreis, Stetten/Rems, Haldenbachtal, auf Mörtel an schattiger Mauer im Wald, 311 m, 48,7764° N, 9,998° E, leg. et det. Schumm 12.05.2010.


Felix Schumm CC BY-SA 4.0
[16492], Germany, Baden-Württemberg, Rems-Murr-Kreis, Stetten/Rems, Haldenbachtal, auf Mörtel an schattiger Mauer im Wald, 311 m, 48,7764° N, 9,998° E, leg. et det. Schumm 12.05.2010.


Felix Schumm CC BY-SA 4.0
[16492], Germany, Baden-Württemberg, Rems-Murr-Kreis, Stetten/Rems, Haldenbachtal, auf Mörtel an schattiger Mauer im Wald, 311 m, 48,7764° N, 9,998° E, leg. et det. Schumm 12.05.2010.


Felix Schumm CC BY-SA 4.0
[16492], Germany, Baden-Württemberg, Rems-Murr-Kreis, Stetten/Rems, Haldenbachtal, auf Mörtel an schattiger Mauer im Wald, 311 m, 48,7764° N, 9,998° E, leg. et det. Schumm 12.05.2010.


Felix Schumm CC BY-SA 4.0
[16492], Germany, Baden-Württemberg, Rems-Murr-Kreis, Stetten/Rems, Haldenbachtal, auf Mörtel an schattiger Mauer im Wald, 311 m, 48,7764° N, 9,998° E, leg. et det. Schumm 12.05.2010.


Felix Schumm CC BY-SA 4.0
[16492], Germany, Baden-Württemberg, Rems-Murr-Kreis, Stetten/Rems, Haldenbachtal, auf Mörtel an schattiger Mauer im Wald, 311 m, 48,7764° N, 9,998° E, leg. et det. Schumm 12.05.2010.