Altitudinal range: from the submediterranean/colline belt (potential vegetation: mixed deciduous forests dominated by Quercus and Carpinus) to the montane belt (potential vegetation: deciduous forests dominated by Fagus sylvatica and closed coniferous forests with Picea abies)
Note: on hard siliceous rocks near the ground in cold-humid habitats, sometimes on walls, mostly below the montane belt; in the study area so far recorded from a single station at the limit of the Southern Pre-Alps (France).
