Altitudinal distribution: mesomediterranean belt (potential vegetation: evergreen broad-leaved forests dominated by Quercus ilex)
Note: on steeply inclined to underhanging surfaces of base-rich or calciferous rocks, sometimes on soil in rock fissures, mostly in warm-dry situations, e.g. in grasslands and garrigues; in the study area only known from the Western Alps (France).
