Altitudinal range: from the mesomediterranean belt (potential vegetation: evergreen broad-leaved forests dominated by Quercus ilex) to the submediterranean/colline belt (potential vegetation: mixed deciduous forests dominated by Quercus and Carpinus)
Note: on calciferous, clayey soil; only known from Dalmatia, France and Piemonte, this species is worthy of further study.