Altitudinal range: from the mesomediterranean belt (potential vegetation: evergreen broad-leaved forests dominated by Quercus ilex) to the montane belt (potential vegetation: deciduous forests dominated by Fagus sylvatica and closed coniferous forests with Picea abies)
Note: on hard siliceous rocks near the shoreline, but sometimes also found at some distance from the coast; in the study area only known from the Western Alps (France); an earlier record from Switzerland (VS) has been excluded, as it is most probably wrong.
