LICHALP

Thelidium papulare (Fr.) Arnold

Syn.: Arthopyrenia sprucei (Bab.) H. Olivier, Polyblastia papularis (Fr.) Servít, Sagedia lariana (A. Massal.) Anzi, Thelidium jurassicum Zschacke, Thelidium larianum A. Massal., Thelidium pyrenophorum sensu A. Massal., Thelidium rubellum A. Massal., Thelidium sprucei (Bab.) Lönnr., Thelidium subpapulare Zschacke, Thelidium umbilicatum Th. Fr., ?Thelidium variabile B. de Lesd., Verrucaria cryptarum Garov., Verrucaria leonina Anzi, Verrucaria papularis Fr., Verrucaria sprucei Bab.; incl. Thelidium papulare (Fr.) Arnold f. fuscum Zschacke

Lichenised.

Substrate: calciferous rocks, intermediate rocks (such as calciferous schists)

Altitudinal range: from the montane belt (potential vegetation: deciduous forests dominated by Fagus sylvatica and closed coniferous forests with Picea abies) to the nival belt (above the lower limit of perennial snow and glaciers)

Note: an arctic-alpine to boreal-montane, circumpolar species of calcareous rocks, with optimum on limestone and dolomite, but also found on calciferous schist and sandstone in upland areas, sometimes growing in temporarily submerged sites along creeks; widespread throughout the Alps.