Altitudinal range: from the submediterranean/colline belt (potential vegetation: mixed deciduous forests dominated by Quercus and Carpinus) to the alpine belt (potential vegetation: treeless Alpine grasslands and tundras, to the lower limit of perennial snow and the equilibrium line of glaciers)
Note: on soil amongst bryophytes and on plant debris on more or less calcareous substrata, apparently with a wide altitudinal range. This taxon deserves further study; in the study area only recorded from a single station in the Eastern Alps (Italy).
