Altitudinal range: from the submediterranean/colline belt (potential vegetation: mixed deciduous forests dominated by Quercus and Carpinus) to the montane belt (potential vegetation: deciduous forests dominated by Fagus sylvatica and closed coniferous forests with Picea abies)
Note: a mild-temperate to tropical species found on smooth bark, especially of Fraxinus and Carpinus, in humid woodlands, with a few records from the Western Alps.
